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Shih, H.-T., P.K.L. Ng, P.J.F. Davie, C.D. Schubart, M. Türkay, R. Naderloo, D.S. Jones, and M.-Y. Liu (2016) Systematics of the family Ocypodidae Rafinesque, 1815 (Crustacea: Brachyura), based on phylogenetic relationships, with a reorganization of subfamily rankings and a review of the taxonomic status of Uca Leach, 1814, sensu lato and its subgenera. Raffles Bulletin of Zoology 64:139–175.

Language: English

Names Appearing in this Publication

Name Used Common Name(s) Where Applied to... Accepted Name Source of Accepted Note(s)
Afruca   text p. 139-141, 143-145, 147-149, 161, 168 citation: Beinlich & von Hagen (2006)Uca [Crane1975] Afruca Computed in part
      citation: Crane (1975)Afruca Afruca Computed  
      citation: Levinton et al. (1996) TBD Computed in part
      citation: Ng et al. (2008) TBD Computed in part
      citation: Rosenberg (2001)Uca [Crane1975] Afruca Computed in part
      citation: Spivak et al. (1991) TBD Computed  
      citation: Sturmbauer et al. (1996) TBD Computed in part
 
Afruca tangeri   text p. 140, 143, 147-149, 168; Figures 6A-B citation: Bott (1973)Uca tangeri Uca tangeri Computed  
      citation: Eydoux (1835)Gelasimus Tangeri Uca tangeri Computed  
      citation: Herklots (1851)Gelasimus perlatus Uca tangeri Computed  
      citation: Holthuis (1979) TBD Computed  
      citation: ICZN (1983)Gelasimus tangeri Uca tangeri Computed  
      citation: Monod (1927)Uca Tangeri var. platydactylus [Manoka Bay, Cameroon] Uca tangeri Computed  
      citation: Monod (1929)Uca tangeri var. matandensis Uca tangeri Computed  
      citation: Rochebrune (1883)Gelasimus cimatodus Uca tangeri Computed  
      citation: Rosenberg (2001)Uca tangeri Uca tangeri Computed in part
      citation: Spivak et al. (1991) TBD Computed  
      specimen: El Puerto de Santa Maria, Cádiz, Spain [NCHUZOOL 13585] Uca tangeri Original  
      specimen: El Puerto de Santa Maria, Cádiz, Spain [NCHUZOOL 14911] Uca tangeri Original  
      specimen: Elmina, Ghana [NCHUZOOL 13654] Uca tangeri Original  
 
Austruca   text p. 139-140, 142, 144, 151-153, 156, 161, 168 citation: Bott (1973)Austruca TBD Computed  
      citation: Crane (1975)Celuca Leptuca Computed in part
      citation: Shih et al. (2013) TBD Computed  
 
Austruca albimana   text p. 142, 153, 168 citation: Beinlich & von Hagen (2006)Uca (Paraleptuca) albimana Uca albimana Computed  
      citation: Kossmann (1877)Gelasimus annulipes var. albimana Uca albimana Computed  
      specimen: Nabq, Sinai, Egypt [NCHUZOOL 13242] Uca albimana Original  
 
Austruca annulipes   text p. 142, 151-153, 168; Figure 8A citation: Beinlich & von Hagen (2006)Uca (Paraleptuca) annulipes Uca annulipes Computed  
      citation: Milne Edwards (1837)Gelasimus annulipes Uca annulipes Computed  
      citation: Verwey (1930)Uca consobrinus Uca annulipes Computed  
      citation: White (1847)Gelasimus porcellanus Uca annulipes Computed  
      location: Tioman, Malaysia Uca annulipes Original  
      specimen: Phuket Island, Thailand [NCHUZOOL 13258] Uca annulipes Original  
 
Austruca bengali   text p. 142, 152-153, 168; Figure 8B citation: Beinlich & von Hagen (2006)Uca (Paraleptuca) bengali Uca bengali Computed  
      citation: Crane (1975)Uca (Celuca) triangularis bengali Uca bengali Computed in part
      location: Selangor, Malaysia Uca bengali Original  
      specimen: Selangor, Malaysia [NCHUZOOL 13575] Uca bengali Original  
 
Austruca cryptica   text p. 140, 153 citation: Naderloo et al. (2010)Uca (Austruca) cryptica Uca cryptica Computed  
 
Austruca iranica   text p. 140, 142, 152-153, 168; Figure 8C citation: Pretzmann (1971)Uca annulipes iranica Uca iranica Computed  
      location: Iran Uca iranica Original  
      specimen: Gav Bandi, Iran [NCHUZOOL 13245] Uca iranica Original  
 
Austruca lactea   text p. 142, 152-153, 168; Figure 8D citation: Beinlich & von Hagen (2006)Uca (Paraleptuca) lactea Uca lactea Computed  
      citation: Milne Edwards (1837)Gelasimus forceps ? Computed  
      citation: de Haan (1833-1850)Ocypode (Gelasimus) lactea Uca lactea Computed  
      location: Taiwan Uca lactea Original  
      specimen: Hong Kong, China [NCHUZOOL 13250] Uca lactea Original  
 
Austruca mjoebergi   text p. 142, 152-153, 168; Figure 8E citation: Beinlich & von Hagen (2006)Uca (Paraleptuca) mjoebergi Uca mjoebergi Computed  
      citation: Rathbun (1924)Uca mjöbergi Uca mjoebergi Computed  
      location: Northern Territory, Australia Uca mjoebergi Original  
      specimen: Bedford Islands, Western Australia, Australia [QM W20253] Uca mjoebergi Original  
 
Austruca occidentalis   text p. 140, 142, 153, 168 citation: Naderloo et al. (2016) TBD Computed  
      specimen: Madagascar [ZRC THH04-30] Uca occidentalis Original  
 
Austruca perplexa   text p. 142, 152-153, 168; Figure 8F citation: Beinlich & von Hagen (2006)Uca (Paraleptuca) perplexa Uca vocans Computed  
      citation: Milne-Edwards (1873)Gelasimus perplexus Uca perplexa Computed in part
      citation: Nobili (1901)Uca annulipes var. orientalis Uca perplexa Computed  
      location: Dongsha Islands, Taiwan Uca perplexa Original  
      specimen: Dulan Forest, Taitung, Taiwan [NTOU] Uca perplexa Original  
      specimen: Ouano Bay, New Caledonia [NCHUZOOL 13573] Uca perplexa Original  
      specimen: Wallis, Wallis and Futuna [NCHUZOOL 14912] Uca perplexa Original  
 
Austruca sindensis   text p. 140, 142, 152-153, 156, 168; Figure 8G citation: Alcock (1900)Gelasimus inversus, var. sindensis Uca sindensis Computed  
      citation: Crane (1975)Uca (Amphiuca) inversa sindensis Uca sindensis Computed  
      citation: Shih et al. (2013) TBD Computed  
      location: Iran Uca sindensis Original  
      specimen: Khor Al-Zubair, Iraq [ZRC 2010.0103] Uca sindensis Original  
      specimen: Qeshm, Iran [NCHUZOOL 13576] Uca sindensis Original  
 
Austruca triangularis   text p. 142, 152-153, 168; Figure 8H citation: Beinlich & von Hagen (2006)Uca (Paraleptuca) triangularis Uca triangularis Computed  
      citation: Milne-Edwards (1873)Gelasimus triangularis Uca triangularis Computed  
      location: Cebu, Philippines Uca triangularis Original  
      specimen: Cebu, Philippines [NCHUZOOL 13574] Uca triangularis Original  
 
Cranuca   text p. 139-140, 142, 144, 151, 153, 155, 161, 169 citation: Beinlich & von Hagen (2006)Cranuca Cranuca Computed in part
      citation: Shih et al. (2013) TBD Computed  
 
Cranuca inversa   text p. 142, 153, 155-156, 169; Figure 9A citation: Beinlich & von Hagen (2006)Uca (Cranuca) inversa Uca inversa Computed  
      citation: Crane (1975)Uca (Amphiuca) inversa inversa Uca inversa Computed  
      citation: Hoffmann (1874)Gelasimus inversus Uca inversa Computed  
      citation: Shih et al. (2013) TBD Computed  
      location: Eastern Africa Uca inversa Original  
      specimen: Dar es Salaam, Tanzania [NCHUZOOL 13255] Uca inversa Original  
      specimen: Gazi, Kenya [MZUF 1024] Uca inversa Original  
      specimen: Sinai, Egypt [TAU SLR 1475] Uca inversa Original  
 
Gelasimus   text p. 139, 142, 144, 149-151, 153, 159, 161, 169 citation: Bott (1973)Latuca TBD Computed  
      citation: Bott (1973)Mesuca TBD Computed  
      citation: Crane (1975)Thalassuca Gelasimus Computed  
      citation: Latreille (1817) TBD Computed  
      citation: Poupin & Juncker (2010) TBD Computed  
 
Gelasimus borealis   text p. 142, 150-151, 169; Figure 7A citation: Chu et al. (2015) TBD Computed  
      citation: Crane (1975)Uca (Thalassuca) vocans borealis Uca borealis Computed  
      location: Taiwan Uca borealis Original  
      specimen: Hong Kong, China [NCHUZOOL 13207] Uca borealis Original  
      specimen: Kinmen, Taiwan [TMCD CHCD853] Uca borealis Original  
 
Gelasimus dampieri   text p. 142, 150-151, 169; Figure 7B citation: Chu et al. (2015) TBD Computed  
      citation: Crane (1975)Uca (Thalassuca) vocans dampieri Uca dampieri Computed  
      location: Northern Territory, Australia Uca dampieri Original  
      specimen: Darwin, Northern Territory, Australia Uca dampieri Original Location mistakenly listed as “New Territory”
 
Gelasimus hesperiae   text p. 142, 150-151, 169; Figure 7C citation: Crane (1975)Uca (Thalassuca) vocans hesperiae Uca hesperiae Computed  
      citation: Kossmann (1877)Gelasimus tetragonon var. spinicarpa Uca hesperiae Computed  
      location: Tanzania Uca hesperiae Original  
      specimen: Madagascar [ZRC THH04-30] Uca hesperiae Original  
      specimen: Mauritius [ZRC THH04-30] Uca hesperiae Original  
      specimen: Mida Creek, Kenya [NCHUZOOL 13172] Uca hesperiae Original  
      specimen: Phuket Island, Thailand [ZRC 2000.1056] Uca hesperiae Original  
 
Gelasimus jocelynae   text p. 140, 142, 150-151, 169; Figure 7D citation: Shih et al. (2010)Uca jocelynae TBD Computed in part
      location: Penghu, Taiwan Uca jocelynae Original  
      specimen: Dulan Forest, Taitung, Taiwan [NTOU] Uca jocelynae Original  
      specimen: Espiritu Santo, Vanuatu [ZRC 2009.0930 [VM53]] Uca jocelynae Original  
      specimen: Panglao, Bohol, Philippines [ZRC 2009.0925 [M9]] Uca jocelynae Original  
      specimen: Papua New Guinea [QM W26812] Uca jocelynae Original  
 
Gelasimus neocultrimana   text p. 142, 150-151, 169; Figure 7E citation: Bott (1973)Mesuca (Latuca) neocultrimana TBD Computed in part
      citation: Crane (1975)Uca (Thalassuca) vocans pacificensis Uca excisa Computed  
      location: Fiji Uca excisa Synonymy  
      specimen: Wallis, Wallis and Futuna [NCHUZOOL 13303] Uca excisa Synonymy  
 
Gelasimus tetragonon   text p. 142, 150-151, 169; Figure 7F citation: Bott (1973)Mesuca (Mesuca) tetragonon Uca tetragonon Computed  
      citation: Crane (1975)Uca (Thalassuca) tetragonon Uca tetragonon Computed in part
      citation: Guérin-Méneville (1829)Gelasimus affinis Uca tetragonon Computed  
      citation: Guérin-Méneville (1829)Gelasimus Duperreyi Uca tetragonon Computed  
      citation: Herbst (1790)Cancer tetragonon Uca tetragonon Computed  
      citation: Hess (1865)Gelasimus variatus Uca tetragonon Computed  
      location: Penghu, Taiwan Uca tetragonon Original  
      specimen: Kenting, Pingtung, Taiwan [TMCD CHCD 526] Uca tetragonon Original  
      specimen: Madagascar [ZRC THH04-17] Uca tetragonon Original Location listed as Sarodrano, Madagascar, which is completely in the interior of the island.
      specimen: Mo'orea, French Polynesia [MNHN] Uca tetragonon Original  
 
Gelasimus vocans   text p. 142, 150-151, 169; Figure 7G citation: Chu et al. (2015) TBD Computed  
      citation: Dana (1851)Gelasimus nitidus Uca excisa Computed  
      citation: Desmarest (1823)Gelasimus Marionis Uca vocans Computed  
      citation: Linnaeus (1758)Cancer vocans [Rumphius1705] Uca vocans Computed  
      citation: Milne Edwards (1837)Gelasimus vocans Uca vocans Computed in part
      citation: Nobili (1906)Uca Marionis var. excisa Uca vocans Computed  
      citation: White (1847)Gelasimus cultrimanus TBD Computed  
      location: Labuan, Malaysia Uca vocans Original  
      specimen: Bohol, Philippines [NCHUZOOL 13667] Uca vocans Original  
      specimen: Hainan Island, China [NCHUZOOL 13182] Uca vocans Original  
 
Gelasimus vomeris   text p. 142, 150-151, 169; Figure 7H citation: Chu et al. (2015) TBD Computed  
      citation: McNeill (1920)Uca marionis, var. vomeris Uca vomeris Computed in part
      location: Queensland, Australia Uca vomeris Original  
      specimen: Boggy Creek, Queensland, Australia [QM W23884] Uca vomeris Original  
 
Leptuca   text p. 139, 141, 143-144, 151, 153-156, 161, 169 citation: Barnwell & Thurman (1984) TBD Computed  
      citation: Beinlich & von Hagen (2006)Leptuca TBD Computed in part
      citation: Bezerra (2012) TBD Computed  
      citation: Bott (1973)Leptuca TBD Computed  
      citation: Bott (1973)Planuca TBD Computed  
      citation: Crane (1975)Celuca Leptuca n part  
      citation: Crane (1975)Boboruca Planuca Computed  
      citation: Levinton et al. (1996) TBD Computed  
      citation: Ng et al. (2008) TBD Computed  
      citation: Rosenberg (2001)Leptuca Leptuca Computed in part
      citation: Shih et al. (2013) TBD Computed  
      citation: Shih et al. (2015) TBD Computed  
      citation: Sturmbauer et al. (1996) TBD Computed  
 
Leptuca batuenta   text p. 153 citation: Crane (1941)Uca batuenta Uca batuenta Computed  
 
Leptuca beebei   text p. 153 citation: Crane (1941)Uca beebei Uca beebei Computed in part
 
Leptuca coloradensis   text p. 153 citation: Rathbun (1893)Gelasimus coloradensis Uca coloradensis Computed  
 
Leptuca crenulata   text p. 153 citation: Lockington (1877)Gelasimus crenulatus Uca crenulata Computed  
      citation: Rathbun (1893)Gelasimus gracilis Uca crenulata Computed  
 
Leptuca cumulanta   text p. 143, 153, 169 citation: Crane (1943)Uca cumulanta Uca cumulanta Computed  
      specimen: Rio de Janeiro, Brazil [NCHUZOOL 13949] Uca cumulanta Original  
 
Leptuca deichmanni   text p. 143, 153, 155, 169; Figure 9B citation: Rathbun (1935)Uca deichmanni Uca deichmanni Computed  
      location: Panama Uca deichmanni Original  
      specimen: Isla Naos, Panama [NCHUZOOL 13583] Uca deichmanni Original  
 
Leptuca dorotheae   text p. 143, 153, 169 citation: von Hagen (1968)Uca dorotheae Uca dorotheae Computed  
      specimen: Rio Tempisque, Costa Rica [ZRC] Uca dorotheae Original  
 
Leptuca festae   text p. 153 citation: Bott (1954)Uca (Minuca) orthomana Uca festae Computed  
      citation: Bott (1954)Uca (Minuca) leptochela Uca festae Computed  
      citation: Bott (1958)Uca (Minuca) leptochela eibli ? Computed  
      citation: Nobili (1901)Uca Festae Uca festae Computed  
      citation: Rathbun (1935)Uca guayaquilensis Uca festae Computed  
 
Leptuca helleri   text p. 153 citation: Rathbun (1902)Uca helleri Uca helleri Computed  
 
Leptuca inaequalis   text p. 153 citation: Rathbun (1935)Uca inaequalis Uca inaequalis Computed  
 
Leptuca latimanus   text p. 153 citation: Rathbun (1893)Gelasimus latimanus Uca latimanus Computed  
 
Leptuca leptodactyla   text p. 143, 153-154, 170 citation: Chace & Hobbs (1969)Uca leptodactyla Uca leptodactyla Computed  
      citation: Crane (1975)Uca (Celuca) [leptodactyla] leptodactyla Uca leptodactyla Computed  
      citation: Ng et al. (2008) TBD Computed  
      citation: Rankin (1898)Uca leptodactyla Uca leptodactyla Computed  
      citation: Rathbun (1898)Uca leptodactyla Uca leptodactyla Computed  
      specimen: Enseada do Mucuripe, Fortaleza, Ceará, Brazil [NCHUZOOL 14915] Uca leptodactyla Original  
      specimen: Pigeon Creek, San Salvador, The Bahamas [ZRC] Uca leptodactyla Original  
      specimen: São Paulo, Brazil [NCHUZOOL 14914] Uca leptodactyla Original  
 
Leptuca limicola   text p. 153 citation: Crane (1941)Uca limicola Uca limicola Computed  
 
Leptuca musica   text p. 153 citation: Rathbun (1915)Uca musica Uca musica Computed in part
 
Leptuca oerstedi   text p. 153 citation: Rathbun (1904)Uca œrstedi Uca oerstedi Computed  
 
Leptuca panacea   text p. 143, 153, 170 citation: Novak & Salmon (1974)Uca panacea Uca panacea Computed in part
      specimen: South Padre Island, Cameron County, Texas, USA [NCHUZOOL 13950] Uca panacea Original  
 
Leptuca pugilator   text p. 143, 153, 170 citation: Bosc (1802)Ocypoda heterochelos Uca tangeri Computed in part
      specimen: Georgetown, South Carolina, USA [NCHUZOOL 13586] Uca pugilator Original  
      specimen: Seahorse Key, Florida, USA [ASIZ] Uca pugilator Original  
 
Leptuca pygmaea   text p. 153-154 citation: Beinlich & von Hagen (2006)Uca (Leptuca) pygmaea Uca pygmaea Computed  
      citation: Crane (1941)Uca pygmaea Uca pygmaea Computed  
 
Leptuca saltitanta   text p. 154 citation: Crane (1941)Uca saltitanta Uca saltitanta Computed  
 
Leptuca speciosa   text p. 143, 154-155, 170; Figure 9C citation: Ives (1891)Gelasimus speciosus Uca speciosa Computed  
      location: The Bahamas Uca speciosa Original  
      specimen: Alligator Point, Florida, USA [NCHUZOOL 13947] Uca speciosa Original  
      specimen: Pigeon Creek, San Salvador, The Bahamas [ZRC] Uca speciosa Original  
 
Leptuca spinicarpa   text p. 143, 154, 170 citation: Rathbun (1900)Uca spinicarpa Uca spinicarpa Computed  
      specimen: Bolivar Peninsula, Texas, USA [ZRC 2009.0295] Uca spinicarpa Original  
      specimen: Money Bayou, Florida, USA [NCHUZOOL 13947] Uca spinicarpa Original  
 
Leptuca stenodactylus   text p. 143, 154, 170 citation: Milne Edwards & Lucas (1843)Gelasimus stenodactylus Uca stenodactylus Computed  
      citation: Smith (1870)Gelasimus gibbosus Uca stenodactylus Computed  
      specimen: El Salvador [SMF 2357] Uca stenodactylus Original  
 
Leptuca subcylindrica   text p. 143, 154, 170 citation: Beinlich & von Hagen (2006)Uca (Leptuca) subcylindrica Uca subcylindrica Computed in part
      citation: Stimpson (1859)Gelasimus subcylindricus Uca subcylindrica Computed  
      specimen: Kingsville, Texas, USA [NCHUZOOL 13952] Uca subcylindrica Original  
 
Leptuca tallanica   text p. 154 citation: von Hagen (1968)Uca tallanica Uca tallanica Computed  
 
Leptuca tenuipedis   text p. 154 citation: Crane (1941)Uca tenuipedis Uca tenuipedis Computed  
 
Leptuca terpsichores   text p. 143, 154-155, 170; Figure 9D citation: Crane (1941)Uca terpsichores Uca terpsichores Computed  
      location: Panama Uca terpsichores Original  
      specimen: Isla Naos, Panama [NCHUZOOL 13582] Uca terpsichores Original  
 
Leptuca thayeri   text p. 143, 154-155, 170; Figure 9E citation: Rathbun (1900)Uca thayeri Uca thayeri Computed  
      citation: Shih et al. (2015) TBD Computed  
      location: Brazil Uca thayeri Original  
      specimen: Anchieta, Espírito Santo, Brazil [NCHUZOOL 13954] Uca thayeri Original  
      specimen: Hutchinson Island, Fort Pierce, Florida, USA [NCHUZOOL 13953] Uca thayeri Original  
 
Leptuca tomentosa   text p. 140, 154 citation: Bott (1954)Uca (Minuca) mertensi Uca tomentosa Computed  
      citation: Crane (1941)Uca tomentosa Uca tomentosa Computed  
      citation: Ng et al. (2008) TBD Computed  
 
Leptuca umbratila   text p. 143, 154-155, 170; Figure 9F citation: Bott (1954)Uca (Minuca) thayeri zilchi Uca umbratila Computed  
      citation: Crane (1941)Uca umbratila Uca umbratila Computed  
      citation: Shih et al. (2015) TBD Computed  
      location: Panama Uca umbratila Original  
      specimen: Diablo, Panama City, Panama [NCHUZOOL 13579] Uca umbratila Original  
 
Leptuca uruguayensis   text p. 143, 154, 171 citation: Nobili (1901)Uca uruguayensis Uca uruguayensis Computed  
      citation: Oliveira (1939)Uca olympioi Uca uruguayensis Computed  
      specimen: Bahía Samborombón, Argentina [NCHUZOOL 13577] Uca uruguayensis Original  
 
Minuca   text p. 139-141, 143-144, 151, 154, 156-157, 161, 171 citation: Barnwell & Thurman (1984) TBD Computed  
      citation: Beinlich & von Hagen (2006)Minuca Minuca Computed in part
      citation: Bezerra (2012) TBD Computed  
      citation: Bott (1954)Minuca Minuca Computed  
      citation: Crane (1975)Minuca Minuca Computed  
      citation: Levinton et al. (1996) TBD Computed  
      citation: Ng et al. (2008) TBD Computed  
      citation: Rosenberg (2001)Minuca Minuca Computed  
      citation: Shih et al. (2013) TBD Computed  
      citation: Shih et al. (2015) TBD Computed  
      citation: Sturmbauer et al. (1996) TBD Computed  
 
Minuca argillicola   text p. 143, 154, 156, 171 citation: Beinlich & von Hagen (2006)Uca (Minuca) argillicola Uca argillicola Computed in part
      citation: Crane (1941)Uca argillicola Uca argillicola Computed  
      specimen: Puerto El Morro, Ecuador [SMF 34737] Uca argillicola Original  
 
Minuca brevifrons   text p. 143, 154-156, 171; Figure 9G citation: Shih et al. (2015) TBD Computed  
      citation: Stimpson (1860)Gelasimus brevifrons Uca brevifrons Computed  
      location: Costa Rica Uca brevifrons Original  
      specimen: San Juanillo, Costa Rica [ZRC 2012.0126] Uca brevifrons Original  
 
Minuca burgersi   text p. 143, 154-156, 171; Figure 9H citation: Coelho (1972) TBD Computed  
      citation: Holthuis (1967)Uca burgersi Uca burgersi Computed in part
      citation: Shih et al. (2015) TBD Computed  
      location: The Bahamas Uca burgersi Original  
      specimen: Brazil [NCHUZOOL 13956] Uca burgersi Original  
      specimen: San Salvador, The Bahamas [ZRC] Uca burgersi Original  
 
Minuca ecuadoriensis   text p. 143, 154, 156, 171 citation: Crane (1943)Uca schmitti Uca ecuadoriensis Computed  
      citation: Landstorfer & Schubart (2010)Uca ecuadoriensis Uca ecuadoriensis Computed  
      citation: Maccagno (1928)Uca ecuadoriensis Uca ecuadoriensis Computed  
      specimen: Puerto El Morro, Ecuador [SMF 34740] Uca ecuadoriensis Original  
 
Minuca galapagensis   text p. 143, 154, 156, 171 citation: Landstorfer & Schubart (2010)Uca galapagensis Uca galapagensis Computed  
      citation: Milne Edwards & Lucas (1843)Gelasimus macrodactylus ? Computed  
      citation: Rathbun (1902)Uca galapagensis Uca galapagensis Computed  
      specimen: Puerto El Morro, Ecuador [SMF 34741] Uca galapagensis Original  
 
Minuca herradurensis   text p. 143, 154, 156-157, 171; Figure 10A citation: Bott (1954)Uca (Minuca) galapagensis herradurensis Uca herradurensis Computed  
      citation: Landstorfer & Schubart (2010)Uca herradurensis Uca herradurensis Computed  
      location: Panama Uca herradurensis Original  
      specimen: Diablo, Panama City, Panama [NCHUZOOL 13580] Uca herradurensis Original  
 
Minuca longisignalis   text p. 143, 154, 156, 171 citation: Salmon & Atsaides (1968)Uca longisignalis Uca longisignalis Computed  
      citation: Shih et al. (2015) TBD Computed  
      specimen: Ingleside Cove, Corpus Christi, Texas, USA [NCHUZOOL 13938] Uca < ongisignalis They suggest this species may be identical to Minuca minax.
 
Minuca marguerita   text p. 154, 156 citation: Thurman (1981)Uca marguerita Uca marguerita Computed in part
 
Minuca minax   text p. 143, 154, 171 citation: Le Conte (1855)Gelasimus minax Uca minax Computed  
      specimen: Chesapeake Bay, Virginia, USA [NCHUZOOL 13939] Uca minax Original  
      specimen: Florida, USA [NCHUZOOL 13957] Uca minax Original  
 
Minuca mordax   text p. 143, 154, 156, 171 citation: Shih et al. (2015) TBD Computed  
      citation: Smith (1870)Gelasimus mordax Uca mordax Computed  
      specimen: São Paulo, Brazil [NCHUZOOL 13940] Uca mordax Original  
 
Minuca osa   text p. 140, 143, 154, 156, 171 citation: Landstorfer & Schubart (2010)Uca osa Uca osa Computed  
      specimen: Golfo Dolce, Costa Rica [UCR 2620-01] Uca osa Original  
 
Minuca pugnax   text p. 143, 154, 171 citation: Rafinesque (1817)Ocypoda pusilla Uca pugnax Computed  
      citation: Smith (1870)Gelasimus pugnax Uca pugnax Computed in part
      specimen: Assateague Island, Maryland, USA [NCHUZOOL 13941] Uca pugnax Original  
 
Minuca rapax   text p. 143, 154, 157, 171; Figure 10B citation: Milne Edwards (1852)Gelasimus palustris Uca vocans Computed in part
      citation: Oliveira (1939)Uca pugnax brasiliensis Uca rapax Computed  
      citation: Smith (1870)Gelasimus rapax Uca rapax Computed  
      location: Brazil Uca rapax Original  
      specimen: Bocas del Toro, Panama [NCHUZOOL 13944] Uca rapax Original  
      specimen: Paraquita Bay, British Virgin Islands [NCHUZOOL 13943] Uca rapax Original  
      specimen: Trelawny, Jamaica [NCHUZOOL 13942] Uca rapax Original  
 
Minuca victoriana   text p. 143, 154, 171 citation: von Hagen (1987)Uca victoriana Uca victoriana Computed  
      specimen: Bahia, Brazil [NCHUZOOL 13945] Uca victoriana Original  
 
Minuca virens   text p. 140-141, 143, 154, 156, 171-172 citation: Rosenberg (2014) TBD Computed  
      citation: Salmon & Atsaides (1968)Uca virens Uca virens Computed  
      citation: Shih et al. (2015) TBD Computed  
      specimen: Ingleside Cove, Corpus Christi, Texas, USA [NCHUZOOL 13584] Uca virens Original  
      specimen: Money Bayou, Florida, USA [NCHUZOOL 13946] Uca virens Original  
 
Minuca vocator   text p. 143, 154, 156, 172 citation: Crane (1943)Uca murifecenta Uca vocator Computed  
      citation: Herbst (1804)Cancer vocator Uca vocator Computed  
      citation: Landstorfer & Schubart (2010)Uca vocator Uca vocator Computed  
      citation: Oliveira (1939)Uca salsisitus Uca rapax Computed  
      citation: von Hagen (1968)Uca lanigera Uca ecuadoriensis Computed  
      specimen: Ceará, Brazil [NCHUZOOL 13948] Uca vocator Original  
      specimen: Trinidad Uca vocator Original  
 
Minuca zacae   text p. 143, 154, 172 citation: Bott (1954)Uca (Minuca) macrodactyla glabromana Uca herradurensis Computed in part
      citation: Crane (1941)Uca zacae Uca zacae Computed  
      specimen: El Salvador [SMF 2104a] Uca zacae Original  
 
Paraleptuca   text p. 139, 142, 144, 151, 153, 156-157, 159, 161, 173 citation: Beinlich & von Hagen (2006)Paraleptuca TBD Computed in part; in part
      citation: Bott (1973)Paraleptuca TBD Computed  
      citation: Crane (1975)Amphiuca Paraleptuca Computed  
      citation: Poupin & Juncker (2010) TBD Computed  
      citation: Shih et al. (2012) TBD Computed  
      citation: Shih et al. (2013)Paraleptuca Paraleptuca Computed  
 
Paraleptuca boninensis   text p. 140, 156 citation: Shih et al. (2013)Uca (Paraleptuca) boninensis TBD Computed in part
 
Paraleptuca chlorophthalmus   text p. 142, 156-157, 173; Figure 10C citation: Beinlich & von Hagen (2006)Uca (Paraleptuca) chlorophthalmus Uca chlorophthalmus Computed  
      citation: Crane (1975)Uca (Amphiuca) chlorophthalmus chlorophthalmus Uca chlorophthalmus Computed in part
      citation: Doflein (1899)Uca amazonensis Uca chlorophthalmus Computed  
      citation: Milne Edwards (1837)Gelasimus chlorophthalmus Uca chlorophthalmus Computed  
      location: Eastern Africa Uca chlorophthalmus Original  
      specimen: Dar es Salaam, Tanzania [NCHUZOOL 13561] Uca chlorophthalmus Original  
      specimen: Malamani, Mayotte [MNHN-IU-2011-5599] Uca chlorophthalmus Original  
      specimen: Malamani, Mayotte [MNHN-IU-2011-5600] Uca chlorophthalmus Original  
 
Paraleptuca crassipes   text p. 142, 156-157, 173; Figure 10D citation: Beinlich & von Hagen (2006)Uca (Paraleptuca) crassipes Uca crassipes Computed  
      citation: Crane (1975)Uca (Amphiuca) chlorophthalmus crassipes Uca crassipes Computed in part
      citation: Milne Edwards (1852)Gelasimus Gaimardi Uca crassipes Computed  
      citation: Milne Edwards (1852)Gelasimus Latreillei Uca crassipes Computed  
      citation: Rathbun (1913)Uca novægunieæ Uca crassipes Computed  
      citation: Stimpson (1858)Gelasimus pulchellus Uca crassipes Computed  
      citation: White (1847)Gelasimus crassipes Uca crassipes Computed  
      location: Dongsha Islands, Taiwan Uca crassipes Original  
      specimen: Ha'apiti, Mo'orea, French Polynesia [NCHUZOOL 13478] Uca crassipes Original  
      specimen: Okinawa Island, Japan [NCHUZOOL 13467] Uca crassipes Original  
 
Paraleptuca splendida   text p. 142, 156-157, 173; Figure 10E citation: Shih et al. (2012) TBD Computed  
      citation: Stimpson (1858)Gelasimus splendidus Uca splendida Computed  
      location: Penghu, Taiwan Uca splendida Original  
      specimen: Cingluo, Penghu, Taiwan [NCHUZOOL 13457] Uca splendida Original  
      specimen: Nha Trang, Vietnam [NCHUZOOL 13561] Uca splendida Original  
      specimen: Tai Tam, Hong Kong, China [NCHUZOOL 13368] Uca splendida Original  
 
Petruca   text p. 139-141, 143-144, 151, 154, 156-157, 161, 173 citation: Shih et al. (2015) TBD Computed  
 
Petruca panamensis   text p. 140, 143, 154, 156-157, 173; Figure 10F citation: Shih et al. (2015) TBD Computed  
      citation: Stimpson (1859)Gelasimus panamensis Uca panamensis Computed  
      location: Costa Rica Uca panamensis Original  
      specimen: Isla Naos, Panama [NCHUZOOL 13581] Uca panamensis Original  
      specimen: Isla Naos, Panama [USNM 1294205] Uca panamensis Original  
      specimen: San Juanillo, Costa Rica [NCHUZOOL 14753] Uca panamensis Original  
      specimen: San Juanillo, Costa Rica [ZRC 2012.0126] Uca panamensis Original  
 
Tubuca   text p. 139, 141-142, 144-145, 151, 157-161, 173 citation: Bott (1973)Tubuca TBD Computed  
      citation: Crane (1975)Australuca Australuca Computed  
      citation: Crane (1975)Deltuca Tubuca Computed  
      citation: Poupin & Juncker (2010) TBD Computed  
 
Tubuca acuta   text p. 142, 159, 173 citation: Stimpson (1858)Gelasimus acutus Uca acuta Computed  
      specimen: Hainan Island, China [NCHUZOOL 13351] Uca acuta Original  
      specimen: Kinmen, Taiwan [NCHUZOOL 13650] Uca acuta Original  
 
Tubuca arcuata   text p. 142, 157, 159, 173; Figure 10G citation: Milne Edwards (1852)Gelasimus brevipes Uca arcuata Computed  
      citation: de Haan (1833-1850)Ocypode (Gelasimus) arcuata Uca arcuata Computed  
      location: Hainan Island, China Uca arcuata Original  
      specimen: Dongzhai Harbour, Hainan Island, China [NCHUZOOL 13363] Uca arcuata Original  
      specimen: Incheon, South Korea [NCHUZOOL 13651] Uca arcuata Original  
 
Tubuca australiae   text p. 140-141, 159 citation: Crane (1975)Uca (Deltuca) demani australiae ? Computed  
      citation: Ng et al. (2008) TBD Computed  
 
Tubuca bellator   text p. 142, 157, 159, 174; Figure 10H citation: White (1847)Gelasimus bellator Uca bellator Computed  
      location: Labuan, Malaysia Uca bellator Original  
      specimen: Labuan, Malaysia [NCHUZOOL 13649] Uca bellator Original  
 
Tubuca capricornis   text p. 142, 159, 173 citation: Crane (1975)Uca (Deltuca) dussumieri capricornis Uca capricornis Computed  
      citation: George & Jones (1982)Uca pavo Uca capricornis Computed  
      specimen: Bedford Islands, Western Australia, Australia [qM W. 20267] Uca capricornis Original  
 
Tubuca coarctata   text p. 142, 158-159, 173-174; Figure 11A citation: Kirk (1880)Gelasimus thomsoni ? Computed  
      citation: Milne Edwards (1852)Gelasimus coarctatus Uca coarctata Computed  
      citation: Nemec (1939)Uca ischnodactylus Uca coarctata Computed  
      citation: Pearse (1912)Uca rathbunæ Uca coarctata Computed  
      citation: Rathbun (1913)Uca mearnsi Uca coarctata Computed  
      location: Taiwan Uca coarctata Original  
      specimen: New Caledonia [QM W29055] Uca coarctata Original  
      specimen: Penghu, Taiwan [NCHUZOOL 13231] Uca coarctata Original  
      specimen: Queensland, Australia [QM W19245] Uca coarctata Original  
      specimen: Queensland, Australia [QM W19920] Uca coarctata Original  
      specimen: Turtle Bay, Queensland, Australia [QM W20132] Uca coarctata Original  
 
Tubuca demani   text p. 142, 159, 174 citation: Ortmann (1897)Uca demani Uca demani Computed in part
      citation: Rathbun (1913)Uca zamboangana Uca demani Computed  
      specimen: Cebu, Philippines [NCHUZOOL 13372] Uca demani Original  
 
Tubuca dussumieri   text p. 142, 158-159, 174; Figure 11B citation: Milne Edwards (1852)Gelasimus Dussumieri Uca dussumieri Computed in part
      citation: Stimpson (1858)Gelasimus dubius Uca coarctata Computed  
      location: Penghu, Taiwan Uca dussumieri Original  
      specimen: Mindanao, Philippines [NCHUZOOL 14922] Uca dussumieri Original  
      specimen: New Caledonia [QM W29054] Uca dussumieri Original  
      specimen: Penghu, Taiwan [NCHUZOOL 14717] Uca dussumieri Original  
 
Tubuca elegans   text p. 142, 158-159, 174; Figure 11C citation: George & Jones (1982)Uca elegans Uca elegans Computed  
      location: Northern Territory, Australia Uca elegans Original Location mistakenly listed as “New Territory”
      specimen: Lacrosse Island, Western Australia, Australia [QM W21038] Uca elegans Original  
 
Tubuca flammula   text p. 159 citation: Crane (1975)Uca (Deltuca) [coarctata] coarctata flammula Uca flammula Computed in part; correct spelling of name
 
Tubuca flamula   text p. 142, 174 specimen: Dampier, Western Australia, Australia [ZRC Aus 010800/13] Uca flammula Original typo in name
      specimen: Kimberley, Western Australia, Australia [QM W20214] Uca flammula Original typo in name
 
Tubuca forcipata   text p. 142, 158-159, 174; Figure 11D citation: Adams & White (1848)Gelasimus forcipatus Uca forcipata Computed  
      citation: Estampador (1937)Uca rubripes ? Computed in part
      citation: Rathbun (1909)Uca manii Uca acuta Computed  
      location: Sarawak, Borneo, Malaysia Uca forcipata Original  
      specimen: Johor, Malaysia [NTOU] Uca forcipata Original  
 
Tubuca hirsutimanus   text p. 159 citation: George & Jones (1982)Uca hirsutimanus Uca hirsutimanus Computed  
 
Tubuca longidigitum   text p. 142, 159, 174 citation: Kingsley (1880)Gelasimus longidigitum Uca longidigitum Computed  
      specimen: Boggy Creek, Queensland, Australia [QM W23884] Uca longidigitum Original  
      specimen: Hervey Bay, Queensland, Australia [QM W19274] Uca longidigitum Original  
      specimen: Redland Bay, Brisbane, Queensland, Australia [NCHUZOOL 13656] Uca longidigitum Original  
 
Tubuca paradussumieri   text p. 142, 159, 174 citation: Bott (1973)Mesuca (Latuca) paradussumieri TBD Computed in part
      citation: Crane (1975)Uca (Deltuca) dussumieri spinata Uca paradussumieri Computed in part
      specimen: Hainan Island, China [NCHUZOOL 13381] Uca paradussumieri Original  
 
Tubuca polita   text p. 158-159; Figure 11E citation: Crane (1975)Uca (Australuca) polita Uca polita Computed  
      location: Northern Territory, Australia Uca polita Original  
 
Tubuca rhizophorae   text p. 142, 158-159, 174; Figure 11F citation: Tweedie (1950)Uca rhizophorae Uca rhizophorae Computed  
      location: Johor, Malaysia Uca rhizophorae Original  
      specimen: Madang, Sabah, Borneo, Malaysia [NCHUZOOL 14924] Uca rhizophorae Original  
      specimen: Mersing, Johor, Malaysia [NCHUZOOL 14923] Uca rhizophorae Original  
 
Tubuca rosea   text p. 142, 158-159, 174; Figure 11G citation: Tweedie (1937)Gelasimus roseus Uca rosea Computed  
      location: Selangor, Malaysia Uca rosea Original  
      specimen: Johor, Malaysia [NTOU] Uca rosea Original  
 
Tubuca seismella   text p. 142, 159, 175 citation: Crane (1975)Uca (Australuca) seismella Uca seismella Computed  
      specimen: West Papua, Indonesia [ZRC 2000.2059] Uca seismella Original  
 
Tubuca signata   text p. 142, 158-159, 175; Figure 11H citation: Hess (1865)Gelasimus signatus Uca signata Computed  
      location: Northern Territory, Australia Uca signata Original Location mistakenly listed as “New Territory”
      specimen: Hucks Landing, Queensland, Australia [QM W19211] Uca signata Original  
 
Tubuca typhoni   text p. 142, 159, 174 citation: Crane (1975)Uca (Deltuca) demani typhoni Uca typhoni Computed  
      specimen: Hainan Island, China [NCHUZOOL 13371] Uca typhoni Original  
 
Tubuca urvillei   text p. 142, 159-160, 174; Figure 12A citation: Milne Edwards (1852)Gelasimus Urvillei Uca urvillei Computed  
      location: Ranong, Thailand Uca urvillei Original  
      specimen: Mumbai, Maharashtra, India [NCHUZOOL 14925] Uca urvillei Original  
      specimen: Poroani, Mayotte [ZRC 1999.1107] Uca urvillei Original  
 
Uca fiddler crabs text p. 139-141, 143-145, 148-149, 151, 161, 175 citation: Agassiz (?)Eurychelus monilifer Uca monilifera Computed  
      citation: Barnwell (1982) TBD Computed  
      citation: Beinlich & von Hagen (2006)Uca Uca    
      citation: Bott (1973)Gelasimus platydactylus Uca major Computed in part
      citation: Bott (1973)Heteruca TBD Computed  
      citation: Chace & Hobbs (1969) Uca    
      citation: Crane (1975)Uca Uca Computed  
      citation: Landstorfer & Schubart (2010) Uca    
      citation: Leach (1814)Uca Uca Computed  
      citation: Levinton et al. (1996) Uca    
      citation: Milne Edwards (1852) Uca    
      citation: Morgan (1923) TBD Computed  
      citation: Naderloo et al. (2010) Uca    
      citation: Ng et al. (2008) Uca    
      citation: Rathbun (1918) Uca    
      citation: Rosenberg (2001)Uca Uca Computed in part
      citation: Salmon (1983) Uca    
      citation: Seba (1758)Uka una Uca tangeri Computed  
      citation: Shih et al. (2009) Uca    
      citation: Shih et al. (2010) Uca    
      citation: Shih et al. (2012) Uca    
      citation: Shih et al. (2013) Uca    
      citation: Shih et al. (2013) Uca    
      citation: Shih (2015) TBD Computed  
      citation: Shih et al. (2015) TBD Computed  
      citation: Sturmbauer et al. (1996) Uca    
      citation: Yamaguchi (1994) TBD Computed  
      citation: Yamaguchi & Henmi (2001) TBD Computed  
      citation: von Hagen (1976) Uca    
 
Uca heteropleura   text p. 148-149; Figure 6C citation: Smith (1870)Gelasimus heteropleurus Uca heteropleura Computed  
      location: Panama Uca heteropleura Original  
 
Uca insignis   text p. 148-149; Figure 6D citation: Gerstaecker (1856)Gelasimus (Acanthoplax) excellens TBD Computed  
      citation: Milne Edwards (1852)Acanthoplax insignis Uca insignis Computed  
      citation: Smith (1870)Gelasimus armatus Uca insignis Computed  
      location: El Salvador Uca insignis Original  
 
Uca intermedia   text p. 149 citation: von Prahl & Toro (1985)Uca (Uca) intermedia Uca intermedia Computed  
 
Uca major   text p. 143, 148-149, 175; Figure 6E citation: Herbst (1782)Cancer vocans major Uca tangeri Computed in part
      citation: Holthuis (1979) TBD Computed  
      citation: ICZN (1983)Cancer vocans major Uca tangeri Computed in part
      citation: Lamarck (1801)Ocypoda heterochelos Uca tangeri Computed  
      citation: Leach (1814)Uca Una Uca tangeri Computed  
      citation: Milne Edwards (1837)Gelasimus platydactylus Uca tangeri Computed in part
      citation: Rosenberg (2001)Uca major Uca tangeri Computed in part
      citation: Schramm (1867)Gelasimus grangeri Uca major Computed  
      citation: Shaw & Nodder (1802)Cancer uka Uca tangeri Computed  
      location: The Bahamas Uca major Original  
      specimen: Pigeon Creek, San Salvador, The Bahamas [ZRC] Uca major Original  
 
Uca maracoani   text p. 143, 148-149, 175; Figure 6F citation: Latreille (1802-1803)Ocypode maracoani Uca maracoani Computed in part
      location: Brazil Uca maracoani Original  
      specimen: Itapissuma, Pernambuco, Brazil [NCHUZOOL 13955] Uca maracoani Original  
 
Uca monilifera   text p. 149 citation: Crane (1975)Uca (Uca) princeps monilifera Uca monilifera Computed  
      citation: Rathbun (1915)Uca monilifera Uca monilifera Computed  
 
Uca ornata   text p. 149 citation: Crane (1975)Uca (Uca) ornata Uca ornata Computed in part
      citation: Smith (1870)Gelasimus ornatus Uca ornata Computed  
      citation: von Hagen (1968)Uca pizarri Uca ornata Computed  
 
Uca princeps   text p. 143, 148-149, 175; Figure 6G citation: Smith (1870)Gelasimus princeps Uca princeps Computed  
      location: Peru Uca princeps Original  
      specimen: Peru [SMF 13164] Uca princeps Original  
 
Uca stylifera   text p. 143, 148-149, 175; Figure 6H citation: Milne Edwards (1852)Gelasimus styliferus Uca stylifera Computed in part
      citation: Smith (1870)Gelasimus heterophthalmus Uca stylifera Computed  
      location: Panama Uca stylifera Original  
      specimen: Base Naval Vasco Nuñez de Balboa, Panama [NCHUZOOL 13578] Uca stylifera Original  
 
Xeruca   text p. 139-140, 142, 144, 151, 159-161, 175 citation: Shih (2015) TBD Computed  
 
Xeruca formosensis   text p. 140, 142, 159-160, 175; Figure 12B citation: Crane (1975)Uca (Thalassuca) formosensis Uca formosensis Computed  
      citation: Rathbun (1921)Uca formosensis Uca formosensis Computed  
      citation: Rosenberg (2001)Uca formosensis Uca formosensis Computed in part
      citation: Shih et al. (1999) TBD Computed  
      citation: Shih (2015) TBD Computed  
      location: Taiwan Uca formosensis Original  
      specimen: Bazhang River, Taiwan [NCHUZOOL 13742] Uca formosensis Original  
      specimen: Cingluo, Penghu, Taiwan [NCHUZOOL 13770] Uca formosensis Original  
      specimen: Qigu, Tainan, Taiwan [NCHUZOOL 13691] Uca formosensis Original