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Yamaguchi, T. (1994) Fiddler crabs of the genus Uca in the collections of three natural history museums in Europe. 1. The specimens held by the Nationaal Natuurhistorisch Museum, Leiden and the National History Museum, London. Calanus. Bulletin of the Aitsu Marine Biological Station 11:151–189.

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Rosenberg (2013), Shih et al. (2010), Shih et al. (2016)