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Layne, J.E., W.J.P. Barnes, and L.M.J. Duncan (2003) Mechanisms of homing in the fiddler crab Uca rapax 2. Information sources and frame of reference for a path integration system. Journal of Experimental Biology 206(24):4425–4442.

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Bingman & Cheng (2005), Kamran & Moore (2015)