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Garth, J.S., and D.P. Abbott (1980) Brachyura: The True Crabs. Pp. 594–630 in Intertidal Invertebrates of California, R.H. Morris, D.P. Abbott and E.C. Haderlie, eds. Stanford, CA: Stanford University Press.

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Hubbard & Dugan (1989), Villalobos Hiriart et al. (1989)